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I suspect those holes were used to locate the cowl and dash subassembly on the body build truck. On the body framing line several major parts including the cowl, floor pan, front body hinge pillar, roof rails, rocker inner, and miscellaneous brackets and reinforcements had to all come together in perfect alignment to be spot welded together.
 

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Those rubber "grommets" are actually plugs. As Farm Boy said, they are more than likely locating holes used during assembly. Not for windshield washer as those are on the vertical surface of the cowl.

The metal plate is used for cowl/air plenum cleanout. All the leaves and crap end up there. The plate can be removed and that area flushed out. Works great!
 
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